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From the link I provided above:
The extended VMware Tools support does two things:
1. It allows virtual machines running older versions of VMware Tools to run in a supported configuration on newer versions of ESXi.
2. It allows virtual machines running newer versions of VMware Tools to run in a supported configuration on older versions of ESX/ESXi.
So, the answer is YES, you can run a VM with VMware Tools from ESXi 5.5 on a ESXi 4.1 without problems.
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Richardson Porto
Senior Infrastructure Specialist
LinkedIn: http://linkedin.com/in/richardsonporto
Richardson Porto
Senior Infrastructure Specialist
LinkedIn: http://linkedin.com/in/richardsonporto